Marine invertebrates exhibit a wide range of modifications to survive in poorly oxygenated waters, including breathing tubes as in mollusc siphons. Fish have gills instead of lungs, although some species of fish, such as the lungfish, have both. Marine mammals (e.g. dolphins, whales, otters, and seals) need to surface periodically to breathe air. (Full article...)
Teleostei (/ˌtɛliˈɒstiaɪ/; Greekteleios "complete" + osteon "bone"), members of which are known as teleosts (/ˈtɛliɒsts,ˈtiːli-/), is, by far, the largest infraclass in the class Actinopterygii, the ray-finned fishes, and contains 96% of all extant species of fish. Teleosts are arranged into about 40 orders and 448 families. Over 26,000 species have been described. Teleosts range from giant oarfish measuring 7.6 m (25 ft) or more, and ocean sunfish weighing over 2 t (2.0 long tons; 2.2 short tons), to the minute male anglerfishPhotocorynus spiniceps, just 6.2 mm (0.24 in) long. Including not only torpedo-shaped fish built for speed, teleosts can be flattened vertically or horizontally, be elongated cylinders or take specialised shapes as in anglerfish and seahorses.
The difference between teleosts and other bony fish lies mainly in their jaw bones; teleosts have a movable premaxilla and corresponding modifications in the jaw musculature which make it possible for them to protrude their jaws outwards from the mouth. This is of great advantage, enabling them to grab prey and draw it into the mouth. In more derived teleosts, the enlarged premaxilla is the main tooth-bearing bone, and the maxilla, which is attached to the lower jaw, acts as a lever, pushing and pulling the premaxilla as the mouth is opened and closed. Other bones further back in the mouth serve to grind and swallow food. Another difference is that the upper and lower lobes of the tail (caudal) fin are about equal in size. The spine ends at the caudal peduncle, distinguishing this group from other fish in which the spine extends into the upper lobe of the tail fin. (Full article...)
Hemiramphidae is a family of fishes that are commonly called halfbeaks, spipe fish or spipefish. They are a geographically widespread and numerically abundant family of epipelagic fish inhabiting warm waters around the world. The halfbeaks are named for their distinctive jaws, in which the lower jaws are significantly longer than the upper jaws. The similar viviparous halfbeaks (family Zenarchopteridae) have often been included in this family.
Seals range in size from the 1 m (3 ft 3 in) and 45 kg (100 lb) Baikal seal to the 5 m (16 ft) and 3,200 kg (7,100 lb) southern elephant seal. Several species exhibit sexual dimorphism. They have streamlined bodies and four limbs that are modified into flippers. Though not as fast in the water as dolphins, seals are more flexible and agile. Otariids primarily use their front limbs to propel themselves through the water, while phocids and walruses primarily use their hind limbs for this purpose. Otariids and walruses have hind limbs that can be pulled under the body and used as legs on land. By comparison, terrestrial locomotion by phocids is more cumbersome. Otariids have visible external ears, while phocids and walruses lack these. Pinnipeds have well-developed senses—their eyesight and hearing are adapted for both air and water, and they have an advanced tactile system in their whiskers or vibrissae. Some species are well adapted for diving to great depths. They have a layer of fat, or blubber, under the skin to keep warm in cold water, and, other than the walrus, all species are covered in fur. (Full article...)
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3D reconstruction of the skull, viewed from the bottom left, exposing the teeth and internal skull roof
Cartorhynchus (meaning "shortened snout") is an extinct genus of earlyichthyosauriformmarine reptile that lived during the Early Triassicepoch, about 248 million years ago. The genus contains a single species, Cartorhynchus lenticarpus, named in 2014 by Ryosuke Motani and colleagues from a single nearly-complete skeleton found near Chaohu, Anhui Province, China. Along with its close relative Sclerocormus, Cartorhynchus was part of a diversification of marine reptiles that occurred suddenly (over about one million years) during the Spathiansubstage, soon after the devastating Permian-Triassic extinction event, but they were subsequently driven to extinction by volcanism and sea level changes by the Middle Triassic.
Measuring about 40 centimetres (16 in) long, Cartorhynchus was a small animal with a lizard-like body and a short torso; it probably swam in an eel-like manner at slow speeds. Its limbs bore extensive cartilage and could bend like flippers, which may have allowed it to walk on land. The most distinctive features of Cartorhynchus were its short, constricted snout, and its multiple rows of molar-like teeth which grew on the inside surface of its jaw bones. These teeth were not discovered until the specimen was subjected to CT scanning. Cartorhynchus likely preyed on hard-shelled invertebrates using suction feeding, although how it exactly used its inward-directed teeth is not yet known. It was one of up to five independent acquisitions of molar-like teeth among ichthyosauriforms. (Full article...)
Marine mammal adaptation to an aquatic lifestyle varies considerably between species. Both cetaceans and sirenians are fully aquatic and therefore are obligate water dwellers. Pinnipeds are semiaquatic; they spend the majority of their time in the water but need to return to land for important activities such as mating, breeding and molting. Sea otters tend to live in kelp forests and estuaries. In contrast, the polar bear is mostly terrestrial and only go into the water on occasions of necessity, and are thus much less adapted to aquatic living. The diets of marine mammals vary considerably as well; some eat zooplankton, others eat fish, squid, shellfish, or seagrass, and a few eat other mammals. While the number of marine mammals is small compared to those found on land, their roles in various ecosystems are large, especially concerning the maintenance of marine ecosystems, through processes including the regulation of prey populations. This role in maintaining ecosystems makes them of particular concern as 23% of marine mammal species are currently threatened. (Full article...)
The ocean sunfish or common mola (Mola mola) is one of the largest bony fish in the world. It is the type species of the genus Mola, and one of five extant species in the family Molidae. It was once misidentified as the heaviest bony fish, which was actually a different and closely related species of sunfish, Mola alexandrini. Adults typically weigh between 247 and 1,000 kg (545 and 2,205 lb). It is native to tropical and temperate waters around the world. It resembles a fish head without a tail, and its main body is flattened laterally. Sunfish can be as tall as they are long when their dorsal and ventral fins are extended.
Many areas of sunfish biology remain poorly understood,[needs update] and various research efforts are underway, including aerial surveys of populations, satellite surveillance using pop-off satellite tags, genetic analysis of tissue samples, and collection of amateur sighting data. (Full article...)
Steller's sea cow (Hydrodamalis gigas) is an extinctsirenian described by Georg Wilhelm Steller in 1741. At that time, it was found only around the Commander Islands in the Bering Sea between Alaska and Russia; its range extended across the North Pacific during the Pleistoceneepoch, and likely contracted to such an extreme degree due to the glacial cycle. It is possible indigenous populations interacted with the animal before Europeans. Steller first encountered it on Vitus Bering's Great Northern Expedition when the crew became shipwrecked on Bering Island. Much of what is known about its behavior comes from Steller's observations on the island, documented in his posthumous publication On the Beasts of the Sea. Within 27 years of its discovery by Europeans, the slow-moving and easily-caught mammal was hunted into extinction for its meat, fat, and hide.
Some 18th-century adults would have reached weights of 8–10 t (8.8–11.0 short tons) and lengths up to 9 m (30 ft). It was a member of the family Dugongidae, of which the 3 m (9.8 ft) long dugong (Dugong dugon) is the sole living member. It had a thicker layer of blubber than other members of the order, an adaptation to the cold waters of its environment. Its tail was forked, like that of whales or dugongs. Lacking true teeth, it had an array of white bristles on its upper lip and two keratinous plates within its mouth for chewing. It fed mainly on kelp, and communicated with sighs and snorting sounds. Steller believed it was a monogamous and social animal living in small family groups and raising its young, similar to modern sirenians. (Full article...)
Jellyfish are mainly free-swimming marine animals with umbrella-shaped bells and trailing tentacles, although a few are anchored to the seabed by stalks rather than being mobile. The bell can pulsate to provide propulsion for efficient locomotion. The tentacles are armed with stinging cells and may be used to capture prey and defend against predators. Jellyfish have a complex life cycle. The medusa is normally the sexual phase, which produces planula larvae. These then disperse widely and enter a sedentary polyp phase which may include asexual budding before reaching sexual maturity. (Full article...)
Bivalvia (/baɪˈvælviə/), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. The class includes the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. Shipworms bore into wood, clay, or stone and live inside these substances.
The shell of a bivalve is composed of calcium carbonate, and consists of two, usually similar, parts called valves. These valves are for feeding and for disposal of waste. These are joined together along one edge (the hinge line) by a flexible ligament that, usually in conjunction with interlocking "teeth" on each of the valves, forms the hinge. This arrangement allows the shell to be opened and closed without the two halves detaching. The shell is typically bilaterally symmetrical, with the hinge lying in the sagittal plane. Adult shell sizes of bivalves vary from fractions of a millimetre to over a metre in length, but the majority of species do not exceed 10 cm (4 in). (Full article...)
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The orca (Orcinus orca), or killer whale, is a toothed whale that is the largest member of the oceanic dolphin family. It is the only extant species in the genus Orcinus. Orcas are recognizable by their black-and-white patterned body. A cosmopolitan species, they are found in diverse marine environments, from Arctic to Antarctic regions to tropical seas.
Orcas are apex predators with a diverse diet. Individual populations often specialize in particular types of prey. This includes a variety of fish, sharks, rays, and marine mammals such as seals and other dolphins and whales. They are highly social; some populations are composed of highly stable matrilineal family groups (pods). Their sophisticated hunting techniques and vocal behaviors, often specific to a particular group and passed along from generation to generation, are considered to be manifestations of animal culture. (Full article...)
Image 1Cnidarians are the simplest animals with cells organised into tissues. Yet the starlet sea anemone contains the same genes as those that form the vertebrate head. (from Marine invertebrates)
Image 2Estuaries occur when rivers flow into a coastal bay or inlet. They are nutrient rich and have a transition zone which moves from freshwater to saltwater. (from Marine habitat)
Image 10Common-enemy graph of Antarctic food web. Potter Cove 2018. Nodes represent basal species and links indirect interactions (shared predators). Node and link widths are proportional to number of shared predators. Node colors represent functional groups. (from Marine food web)
Image 13Phylogenetic and symbiogenetic tree of living organisms, showing a view of the origins of eukaryotes and prokaryotes (from Marine fungi)
Image 14Food web structure in the euphotic zone. The linear food chain large phytoplankton-herbivore-predator (on the left with red arrow connections) has fewer levels than one with small phytoplankton at the base. The microbial loop refers to the flow from the dissolved organic carbon (DOC) via heterotrophic bacteria (Het. Bac.) and microzooplankton to predatory zooplankton (on the right with black solid arrows). Viruses play a major role in the mortality of phytoplankton and heterotrophic bacteria, and recycle organic carbon back to the DOC pool. Other sources of dissolved organic carbon (also dashed black arrows) includes exudation, sloppy feeding, etc. Particulate detritus pools and fluxes are not shown for simplicity. (from Marine food web)
Image 16An in situ perspective of a deep pelagic food web derived from ROV-based observations of feeding, as represented by 20 broad taxonomic groupings. The linkages between predator to prey are coloured according to predator group origin, and loops indicate within-group feeding. The thickness of the lines or edges connecting food web components is scaled to the log of the number of unique ROV feeding observations across the years 1991–2016 between the two groups of animals. The different groups have eight colour-coded types according to main animal types as indicated by the legend and defined here: red, cephalopods; orange, crustaceans; light green, fish; dark green, medusa; purple, siphonophores; blue, ctenophores and grey, all other animals. In this plot, the vertical axis does not correspond to trophic level, because this metric is not readily estimated for all members. (from Marine food web)
Image 17
Bacterioplankton and the pelagic marine food web
Solar radiation can have positive (+) or negative (−) effects resulting in increases or decreases in the heterotrophic activity of bacterioplankton. (from Marine prokaryotes)
Image 18Antarctic marine food web. Potter Cove 2018. Vertical position indicates trophic level and node widths are proportional to total degree (in and out). Node colors represent functional groups. (from Marine food web)
Image 19A protected sea turtle area that warns of fines and imprisonment on a beach in Miami, Florida. (from Marine conservation)
Image 20Lampreys are often parasitic and have a toothed, funnel-like sucking mouth (from Marine vertebrate)
Image 22Coral reefs provide marine habitats for tube sponges, which in turn become marine habitats for fishes (from Marine habitat)
Image 23Dickinsonia may be the earliest animal. They appear in the fossil record 571 million to 541 million years ago. (from Marine invertebrates)
Image 24This timeline contains clickable links
Image 25Anthropogenic stressors to marine species threatened with extinction (from Marine food web)
Image 26Scanning electron micrograph of a strain of Roseobacter, a widespread and important genus of marine bacteria. For scale, the membrane pore size is 0.2 μm in diameter. (from Marine prokaryotes)
Mycoloop links between phytoplankton and zooplankton
Chytrid‐mediated trophic links between phytoplankton and zooplankton (mycoloop). While small phytoplankton species can be grazed upon by zooplankton, large phytoplankton species constitute poorly edible or even inedible prey. Chytrid infections on large phytoplankton can induce changes in palatability, as a result of host aggregation (reduced edibility) or mechanistic fragmentation of cells or filaments (increased palatability). First, chytrid parasites extract and repack nutrients and energy from their hosts in form of readily edible zoospores. Second, infected and fragmented hosts including attached sporangia can also be ingested by grazers (i.e. concomitant predation). (from Marine fungi)
Image 34The deep sea amphipodEurythenes plasticus, named after microplastics found in its body, demonstrating plastic pollution affects marine habitats even 6000m below sea level. (from Marine habitat)
Image 36Ocean surface chlorophyll concentrations in October 2019. The concentration of chlorophyll can be used as a proxy to indicate how many phytoplankton are present. Thus on this global map green indicates where a lot of phytoplankton are present, while blue indicates where few phytoplankton are present. – NASA Earth Observatory 2019. (from Marine food web)
Image 37Oceanic pelagic food web showing energy flow from micronekton to top predators. Line thickness is scaled to the proportion in the diet. (from Marine food web)
Estimates of microbial species counts in the three domains of life
Bacteria are the oldest and most biodiverse group, followed by Archaea and Fungi (the most recent groups). In 1998, before awareness of the extent of microbial life had gotten underway, Robert M. May estimated there were 3 million species of living organisms on the planet. But in 2016, Locey and Lennon estimated the number of microorganism species could be as high as 1 trillion. (from Marine prokaryotes)
Image 44Elevation-area graph showing the proportion of land area at given heights and the proportion of ocean area at given depths (from Marine habitat)
Image 46Phylogenetic and symbiogenetic tree of living organisms, showing a view of the origins of eukaryotes and prokaryotes (from Marine prokaryotes)
Image 47This algae bloom occupies sunlit epipelagic waters off the southern coast of England. The algae are maybe feeding on nutrients from land runoff or upwellings at the edge of the continental shelf. (from Marine habitat)
Image 48In the open ocean, sunlit surface epipelagic waters get enough light for photosynthesis, but there are often not enough nutrients. As a result, large areas contain little life apart from migrating animals. (from Marine habitat)
Image 49Schematic representation of the changes in abundance between trophic groups in a temperate rocky reef ecosystem. (a) Interactions at equilibrium. (b) Trophic cascade following disturbance. In this case, the otter is the dominant predator and the macroalgae are kelp. Arrows with positive (green, +) signs indicate positive effects on abundance while those with negative (red, -) indicate negative effects on abundance. The size of the bubbles represents the change in population abundance and associated altered interaction strength following disturbance. (from Marine food web)
Image 50Only 29 percent of the world surface is land. The rest is ocean, home to the marine habitats. The oceans are nearly four kilometres deep on average and are fringed with coastlines that run for nearly 380,000 kilometres.
Image 52640 μm microplastic found in the deep sea amphipod Eurythenes plasticus (from Marine habitat)
Image 53A 2016 metagenomic representation of the tree of life using ribosomal protein sequences. The tree includes 92 named bacterial phyla, 26 archaeal phyla and five eukaryotic supergroups. Major lineages are assigned arbitrary colours and named in italics with well-characterized lineage names. Lineages lacking an isolated representative are highlighted with non-italicized names and red dots. (from Marine prokaryotes)
Image 57Sea ice food web and the microbial loop. AAnP = aerobic anaerobic phototroph, DOC = dissolved organic carbon, DOM = dissolved organic matter, POC = particulate organic carbon, PR = proteorhodopsins. (from Marine food web)
Image 58Some representative ocean animal life (not drawn to scale) within their approximate depth-defined ecological habitats. Marine microorganisms exist on the surfaces and within the tissues and organs of the diverse life inhabiting the ocean, across all ocean habitats. (from Marine habitat)
Image 60Cycling of marine phytoplankton. Phytoplankton live in the photic zone of the ocean, where photosynthesis is possible. During photosynthesis, they assimilate carbon dioxide and release oxygen. If solar radiation is too high, phytoplankton may fall victim to photodegradation. For growth, phytoplankton cells depend on nutrients, which enter the ocean by rivers, continental weathering, and glacial ice meltwater on the poles. Phytoplankton release dissolved organic carbon (DOC) into the ocean. Since phytoplankton are the basis of marine food webs, they serve as prey for zooplankton, fish larvae and other heterotrophic organisms. They can also be degraded by bacteria or by viral lysis. Although some phytoplankton cells, such as dinoflagellates, are able to migrate vertically, they are still incapable of actively moving against currents, so they slowly sink and ultimately fertilize the seafloor with dead cells and detritus. (from Marine food web)
Image 61Sponges have no nervous, digestive or circulatory system (from Marine invertebrates)
Image 62
Model of the energy generating mechanism in marine bacteria
(1) When sunlight strikes a rhodopsin molecule (2) it changes its configuration so a proton is expelled from the cell (3) the chemical potential causes the proton to flow back to the cell (4) thus generating energy (5) in the form of adenosine triphosphate. (from Marine prokaryotes)
Image 69Waves and currents shape the intertidal shoreline, eroding the softer rocks and transporting and grading loose particles into shingles, sand or mud (from Marine habitat)
Image 74The Ocean Cleanup is one of many organizations working toward marine conservation such at this interceptor vessel that prevents plastic from entering the ocean. (from Marine conservation)
Image 75Chytrid parasites of marine diatoms. (A) Chytrid sporangia on Pleurosigma sp. The white arrow indicates the operculate discharge pore. (B) Rhizoids (white arrow) extending into diatom host. (C) Chlorophyll aggregates localized to infection sites (white arrows). (D and E) Single hosts bearing multiple zoosporangia at different stages of development. The white arrow in panel E highlights branching rhizoids. (F) Endobiotic chytrid-like sporangia within diatom frustule. Bars = 10 μm. (from Marine fungi)
Image 77Reconstruction of an ammonite, a highly successful early cephalopod that first appeared in the Devonian (about 400 mya). They became extinct during the same extinction event that killed the land dinosaurs (about 66 mya). (from Marine invertebrates)
Image 86Whales were close to extinction until legislation was put in place. (from Marine conservation)
Image 87On average there are more than one million microbial cells in every drop of seawater, and their collective metabolisms not only recycle nutrients that can then be used by larger organisms but also catalyze key chemical transformations that maintain Earth's habitability. (from Marine food web)
Image 101Phylogenetic tree representing bacterial OTUs from clone libraries and next-generation sequencing. OTUs from next-generation sequencing are displayed if the OTU contained more than two sequences in the unrarefied OTU table (3626 OTUs). (from Marine prokaryotes)
Image 103Sandy shores provide shifting homes to many species (from Marine habitat)
Image 104Biomass pyramids. Compared to terrestrial biomass pyramids, aquatic pyramids are generally inverted at the base. (from Marine food web)
Image 105Conceptual diagram of faunal community structure and food-web patterns along fluid-flux gradients within Guaymas seep and vent ecosystems. (from Marine food web)
Image 106A microbial mat encrusted with iron oxide on the flank of a seamount can harbour microbial communities dominated by the iron-oxidizing Zetaproteobacteria (from Marine prokaryotes)
Parasitic chytrids can transfer material from large inedible phytoplankton to zooplankton. Chytrids zoospores are excellent food for zooplankton in terms of size (2–5 μm in diameter), shape, nutritional quality (rich in polyunsaturated fatty acids and cholesterols). Large colonies of host phytoplankton may also be fragmented by chytrid infections and become edible to zooplankton. (from Marine fungi)
Image 111Conference events, such as the events hosted by the United Nations, help to bring together many stakeholders for awareness and action. (from Marine conservation)
Image 113Archaea were initially viewed as extremophiles living in harsh environments, such as the yellow archaea pictured here in a hot spring, but they have since been found in a much broader range of habitats. (from Marine prokaryotes)
Image 114Some lobe-finned fishes, like the extinct Tiktaalik, developed limb-like fins that could take them onto land (from Marine vertebrate)
Image 115Ernst Haeckel's 96th plate, showing some marine invertebrates. Marine invertebrates have a large variety of body plans, which are currently categorised into over 30 phyla. (from Marine invertebrates)
Image 118The pelagic food web, showing the central involvement of marine microorganisms in how the ocean imports nutrients from and then exports them back to the atmosphere and ocean floor (from Marine food web)
Image 119
Different bacteria shapes (cocci, rods and spirochetes) and their sizes compared with the width of a human hair. A few bacteria are comma-shaped (vibrio). Archaea have similar shapes, though the archaeon Haloquadratum is flat and square.
The unit μm is a measurement of length, the micrometer, equal to 1/1,000 of a millimeter
Image 122Topological positions versus mobility: (A) bottom-up groups (sessile and drifters), (B) groups at the top of the food web. Phyto, phytoplankton; MacroAlga, macroalgae; Proto, pelagic protozoa; Crus, Crustacea; PelBact, pelagic bacteria; Echino, Echinoderms; Amph, Amphipods; HerbFish, herbivorous fish; Zoopl, zooplankton; SuspFeed, suspension feeders; Polych, polychaetes; Mugil, Mugilidae; Gastropod, gastropods; Blenny, omnivorous blennies; Decapod, decapods; Dpunt, Diplodus puntazzo; Macropl, macroplankton; PlFish, planktivorous fish; Cephalopod, cephalopods; Mcarni, macrocarnivorous fish; Pisc, piscivorous fish; Bird, seabirds; InvFeed1 through InvFeed4, benthic invertebrate feeders. (from Marine food web)
Image 123Jellyfish are easy to capture and digest and may be more important as food sources than was previously thought. (from Marine food web)
Image 124The distribution of anthropogenic stressors faced by marine species threatened with extinction in various marine regions of the world. Numbers in the pie charts indicate the percentage contribution of an anthropogenic stressors' impact in a specific marine region. (from Marine food web)
Image 21Ecosystem services delivered by epibenthicbivalve reefs. Reefs provide coastal protection through erosion control and shoreline stabilization, and modify the physical landscape by ecosystem engineering, thereby providing habitat for species by facilitative interactions with other habitats such as tidal flat benthic communities, seagrasses and marshes. (from Marine ecosystem)
Image 22Some lobe-finned fishes, like the extinct Tiktaalik, developed limb-like fins that could take them onto land (from Marine vertebrate)
... Some sharks are so flexible, they can bend right around and touch their tails with their snouts.
... the Orca, is the fastest swimmer of all the cetaceans and can reach speeds of more than 50km/h while hunting.
... the Humpback Whales song is produced by them forcing air through their massive nasal cavities
... the Beluga whale is also known as the Sea Canary on account of its high-pitched squeaks, squeals, and whistles.
... on average, a whale or dolphin will eat four to five percent of its body weight in food per day. That means that a 100 ton blue whale will eat almost five tons of krill per day, or that a 200kg bottlenose dolphin will eat 10kg of fish per day!
... in spite of their enormous mass, baleen whales are capable of leaping completely out of the water, particularly the Humpback Whale.
Schoolingbigeye trevally. In biology, any group of fish that stay together for social reasons are said to be shoaling and if the group is swimming in the same direction in a coordinated manner, they are said to be schooling.
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![Image 63Pennate diatom from an Arctic meltpond, infected with two chytrid-like [zoo-]sporangium fungal pathogens (in false-colour red). Scale bar = 10 μm. (from Marine food web)](/wiki/File:Pennate_diatom_infected_with_two_chytrid-like_fungal_pathogens.png)
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